Skull Name: Emperor (Mission 15: Escape) - Kill 3 Scarabs - Head the to Very North of the Map and its dead Center of all the flood Skull Name: Cowbell (Mission 3: Relic Interior) - Kill 45 Hunters - take a Grizzly, Anders, Forge and Your Marines Back to where they were held up on the central pad Skull Name: Catch (Mission 10: Shield World) - Kill 350 Swarms - get at least 2 Hornets and fly East of you base past the First Tower to a Plateau with covered with Flood Eggs it is in the center Skull Name: Bountiful Harvest (Mission 14: Reactor) - Kill 20 Vampires - head to the Second Covenant Base and its in the Far corner at the bottom of the next ramp Skull Name: Boomstick (Mission 12: Repairs) - Kill 12 Spirit Transports - take a Hawk to the Lower 2 Airlocks and it its right up the Little ramp in the Air Mission 15: Escape - The box is at the north edge of the central circle shield thing. Mission 14: Reactor - The box is to the very south west of the map on a cliff ledge which can be accessed from the first elevation, where the first Covenant base is. There are some small ramps which may allow for infantry to get to the box but is a lot easier to just use air units. Mission 13: Beachhead - The box is on the far north east of the map at the top of a cliff. It's on the very west side of the ship under the last gun turret near the power generator.
Mission 12: Repairs - This box is also on the section of ship inaccessible by ground units. It attached to the hull so it may be hard to see. It is on the North West side of the ship where the edge section starts to get wider. Mission 11: Cleansing - The box is on one of the ship sections off to the side which are inaccessible by ground units. Mission 10: Shield World - The box is beside the middle set of units that need evacuation. Mission 9: The Flood - The box is at the west most point of the map in a corner near the bordering mountains.
The box is in the middle of a plateau surrounded by trees. Mission 8: Anders' Signal - Go as far east as possible on the map. Mission 7: Scarab - The box is at the furthest point north east of the map beside a bunch of supply crates. (To reach the island you have to deploy the first two rhino tanks then use the Transport ability to get units there) Mission 6: Dome of Light - Other island where you have to place the third rhino tank, there is an underground entrance south west of the rhino site, the box is in front of this entrance. You need to go down some steps to reach it. The box is to the right of the one near the center of the map. Mission 5: Arcadia Outskirts - There are two capturable reactors on the map. (If you don’t know where the west base location is, just wait until you can build a base after the #3 transport is destroyed) Mission 4: Arcadia City - The box is directly north of the West base location right beside a sky bridge. The box is right before the crashed Pelican. Mission 3: Relic Interior - Right from the start head down the ramp on your right and follow the path all the way to the end. The box is behind this shield (You need infantry to reach it) Mission 2: Relic Approach - North of the map there is an energy shield near the Covenant air pads and Methane Tanks. Go down this ramp and the black box will be right between it and the bridge. Mission 1: Alpha Base - There is a small ramp by the bridge right before you rescue the last set of marines. If all flood monsters were kill when this happened, you will get the achievement during the cinematic. When all 5 are dead, destroy the flood structure marked on your map.
It will be in between the crashed pelican and main base. On your way to each monster, make sure to kill the stalks as killing 20 will unlock the skull. Note that although hornets cannot be touched by most flood, there are still some flying units and tentacles that can kill hornets, so rebuild them as soon as possible. Send each group directly to a flood monster and kill each one until all five are dead at the same time.(They will eventually regenerate so move quick and leave 1 hornet behind to keep it down). Split the remaining hornets into 2-3 groups of about 5 each. Keep 3-4 hornets at your base for protection while building turrets.
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Build up your resources so you can build a large amount of hornets (remember to upgrade them when possible). To beat this level with the achievement is easy.